Lack of data about fish ages
Unlike a few developed countries in fisheries mgt, most countries have not identified ages of fish caught by fisheries and surveys. As a result, we cannot apply an age-structured model (e.g., SCAA).
An alternative
It is relatively easy and much less costly to collect data on fish body sizes (e.g., lengths, weights).
Illustration: Korean chub mackerel (Scomber japonicus) population Â
Longevity: about six years
Spatial distribution
Yields from a purse seine fishery and its CPUE in 1996-2017
Body lengths from 2000-2017; Â Body weights from 2005-2017
To develop a length-based model from scratch (e.g., in ADMB), given the available data about the chub mackerel population. (Completed)
To compare our model in performance with Multifan-CL (or SS). (Ongoing)
Precursors to our model
Cohen and Fishman (1980), Deriso and Parma (1988), and Quinn et al. (1998)
Our model
We extended and modified Quinn’s model from scratch, implementing our model in ADMB as opposed to using existing software such as Multifan-CL.
Cf. Multifan-CL: Schnute and Fournier (1980), Fournier et al. (1990, 1998).
Approach with a mixed effect model
LIME (length-based integrated mixed effects method) (Rudd and Thorn 2017) is attractive (e.g., TMB). But it requires many parameters to be known, which include parameters in length-at-age relationship, natural mortality, parameters in a selectity function, etc.
Then, \(P_{y+1, a+1}(L) => P_{y+1, a+1}(x)\), and steps of (2) ~ (4) are repeated over imaginary ages, or along an imaginary cohort.
where
\(x_{a+1} = h(L_\infty, \color{red}\rho, x_a) + \varepsilon\),  and  \(\varepsilon \text{ ~ } N(0,\color{red}{\sigma^2_x})\)
Year- and length- fishing mortality: Â \(F_{y}(x)=\text{sel}\cdot F_{y} = \text{logistic}(x, \color{red} \gamma,\color{red} {L_{0.5}})\cdot\color{red} q\cdot \text{effort}_y\)
           Â
The number of age-\(a\) fish at length \(x\) after mortality: Â \(N_{y,a,Z}(x) = N_{y,a} \cdot P_{y,a,Z}(x)\)
Catch at age \(a\) and length \(x\): Â \(C_{y,a}(x)=N_{y,a} \cdot P_{y,a}(x) \cdot\frac{F_y(x)}{Z_y(x)}\cdot (1-\text{exp}(-Z_y(x)))\)
Biomass and yield at age \(a\): Â \[B_{y,a}=\sum_{x}^{} N_{y,a}(x)\cdot W(x), \; \; \text{and} \; \;
Y_{y,a}=\sum_{x}^{} C_{y,a}(x)\cdot W(x)\] Â Â Â Â Â Â Â Â Â Â Â Â Â Â where \(W(x)\) = body weight at length \(x\).
\(q\): catchability in the fishing mortality function
\(\gamma\) and \(L_{0.5}\): two parameters in the selection (logistic) equation
\(\color{red}\rho\) and \(\sigma^2_x\): two parameters in the growth equations (Cohen and Fishman; LVB)
\(\mathbf N_1\): 22 annual abundances at recruitment (i.e., in 22 years).
\(M\): natural mortality was not treated as a free parameter but its estimate was found with a sensitivity analysis.
\(\mu_1\) of 18 \(cm\), and \(\sigma^2_1\) of 2.10 \({cm}^2\):
The mean and variance of body lengths of recruitment (i.e., age-1 fish).
\(\beta\) of 3.425, and \(\alpha\) of 0.003 \(gram/cm^{3.425}\):
Parameters in the relationship between body weight and length (i.e., \(W = \alpha \cdot L^\beta\))
\(L_\infty\) of 51.94 \(cm\):
The possible maximum of body length.
\(\mathbf m_y\), length frequency data in year, \(y\): Â Â \(\mathbf m_y\) ~ Multinomial(\(n_y\), \(\mathbf o_j\))
\(Y_y\), yield data in year, \(y\): Â Â \(\text{log} Y_y\) ~ \(N(\mu_{\text{log}Y_y}\), \(\sigma^2_{\text{log}Y})\)
\(\therefore Obj = -1.0\cdot \bigl[D_1 \cdot \text{log}L(\boldsymbol\theta|\mathbf m_y) + D_2 \cdot \text{log}L(\boldsymbol\theta | \text{log}Y_y) \bigr]\)
where \(D_1\) and \(D_2\) are data weights.
It turned out that \(D_1\) = 0.05, and \(D_2\) = 10.0.
1st work was completed but we want to implement process errors in growth parameters to be fully free of equilibrium assumptions.
Comparision between our model and Multifan-CL
If one of you is familiar with running the Multifan-CL (or SS), we would be able to quickly collaborate on the comparision. Contact email: shyunuw@gmail.com
Late Dr. Terry Quinn’s visit at May 2016
Financial support: National Research Foundation of Korea
Data: (Korea) National Institute of Fisheries Sciences
Map: Doyul Kim
Mackerel image: https://en.wikipedia.org/wiki/Chub_mackerel
Thank you.
Supplementary materials